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Ferns and Horsetails (Monilophytes)
Clubmosses and Quillworts (Lycophytes)
Polystichum lonchitis  •  Holly Fern
Polystichum lonchitis (L.) Roth; Holly Fern
Holly Fern, Western Scotland, early August
Single frond. Western Scotland (Argyll), early July Single frond, underside showing sori. Western Scotland (Argyll), early July
Mid section of frond, underside showing sori. Western Scotland (Argyll), early August Underside of pinnae showing sori, some sporangia approaching ripeness. Western Scotland (Argyll), early August
Detail of frond showing pinnae. Yorkshire, late July Yorkshire, late July
Yorkshire, late July In limestone pavement. Yorkshire, late July
Western Scotland (Argyll), early August Western Scotland (Argyll), early July
Western Scotland (Argyll), early July Western Scotland (Argyll), early August
Western Scotland (Argyll), early August An unusual conjunction of Holly Fern, Green Spleenwort and Hard Fern: peaty soil meets the edge of limestone pavement, Yorkshire
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